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1 ehavior, with ramifications beyond bacterial swarming.
2 low viscosity that normally impede wild-type swarming.
3 stimulating biofilm formation and repressing swarming.
4 rs, only one of which can provide torque for swarming.
5 liL cells also exhibit temperature-sensitive swarming.
6 t a particular environment is permissive for swarming.
7 agella as they colonize agar surfaces during swarming.
8 d, a means for surface colonization known as swarming.
9 ation of environmental signals that regulate swarming.
10 ability to inducing activity with respect to swarming.
11 his is a means for colony expansion known as swarming.
12 d that protein synthesis is not required for swarming.
13 h P. mirabilis prepares for the next wave of swarming.
14 h is instrumental for coordinated motion and swarming.
15 scriminate kin from nonkin in the context of swarming, a cooperative multicellular behavior.
16         We delve into Pseudomonas aeruginosa swarming, a phenomenon where billions of bacteria move c
17 vioral rules underlying both aggregation and swarming-a dynamic phenotype only observed at longer tim
18 ferent drying conditions along with the anti-swarming activity against Citrobacter rodentium.
19                                 Natural anti-swarming agents from food waste may have promising poten
20  upregulate flagellar gene expression during swarming, also do not increase flagellar numbers per mum
21 ach colony is to produce all drones prior to swarming, an impossible solution on a population scale b
22 Deletion of MXAN_4832 causes defects in both swarming and aggregation related to cell motility and th
23 e upregulation of VLA3 to support neutrophil swarming and aggregation.
24 n of ICAM1, CD11b, PD-L1 as well as enhanced swarming and aggregation.
25                                    Bacterial swarming and biofilm formation are collective multicellu
26                    Iron availability affects swarming and biofilm formation in various bacterial spec
27 ever, how bacteria sense iron and coordinate swarming and biofilm formation remains unclear.
28 mately play a role in surface-sensing during swarming and biofilm formation.
29 he cylic di-GMP signaling messenger produced swarming and biofilm phenotypes similar to those obtaine
30 ias, which likely accounts for their reduced-swarming and delayed-development phenotypes.
31 uency to support directional motility during swarming and fruiting body formation.
32 sponse characterized by meningeal neutrophil swarming and microglial reconstitution of the damaged gl
33 timulating factors (GCSF and GM-CSF) enhance swarming and neutrophil ability to restrict fungal growt
34                                              Swarming and swimming motility of bacteria has been stud
35 med CalR was identified and shown to repress swarming and T3SS1 gene expression.
36 the physical mechanisms underlying bacterial swarming and the balance between individual and collecti
37 y, we used a computational model to simulate swarming and to probe for individual cell behavior that
38 laP and decreased both putrescine stimulated swarming and urothelial cell invasion in a speA mutant.
39  in Vibrio parahaemolyticus, in part because swarming and virulence factors--the hallmarks of the org
40   In contrast, tan variants are deficient in swarming (+) and persist beyond stationary phase.
41            Yellow variants are proficient at swarming (++) and tend to lyse in liquid during stationa
42           The bioactivity (antioxidant, anti-swarming) and phenolics content was significantly higher
43       RNA was extracted from broth-cultured, swarming, and consolidation-phase cells to assess transc
44                     SscS plays a role during swarming, and mutants lacking this chemoreceptor swarm f
45 luence, prey-predator dynamics, flocking and swarming, and phototaxis in cell dynamics.
46 is study provides detailed insights into the swarming architecture and dynamics of Vibrio alginolytic
47                                     These co-swarming assays further demonstrated that DeltagacA muta
48 tion of motility as measured in swimming and swarming assays.
49           Importantly, choline also enhances swarming-associated colony expansion of P. mirabilis und
50 oss of fliL also results in an inhibition of swarming at <30 degrees C.
51                                   We studied swarming at the cellular level using a combination of la
52 find nutrients and avoid toxic environments, swarming bacteria appear to suppress chemotaxis and to u
53                                              Swarming bacteria can be further subdivided by their req
54                                              Swarming bacteria move on agar surfaces in groups, using
55                                              Swarming bacteria use kin discrimination to preferential
56 f the master flagellar regulators in diverse swarming bacteria.
57 gorithms are presented for the prediction of swarming, based on vibration data recorded using acceler
58 e biosurfactants constitutively we show that swarming becomes cheatable: a non-producing strain rapid
59 ive behavior (0.30) but almost identical for swarming behavior (0.45) compared to corresponding singl
60 ly applicable method for analyzing bacterial swarming behavior in two and three dimensions with both
61 nonmotile cells were sufficient to block the swarming behavior of a large gliding-proficient populati
62  genetic correlation between honey yield and swarming behavior was moderate (0.41).
63 ney production) and threshold (defensive and swarming behavior) single-trait models; estimated geneti
64 energy-harvesting, environmentally triggered swarming behavior, and magnetic control of the new Janus
65 d, 0.36 for defensive behavior, and 0.34 for swarming behavior.
66  correlation was found between defensive and swarming behaviors (0.62).
67  4003 records for honey yield, defensive and swarming behaviors of Italian honey bee queens produced
68  POC originating from Antarctic krill, whose swarming behaviour could result in a major conduit of ca
69 xis and cluster formation reminiscent of the swarming behaviour of insects.
70 duces coordinated, self-organized neutrophil-swarming behaviour that isolates the wound or infectious
71 ems (alternative sigma factors, sporulation, swarming, biofilm formation, stochastic cell fate switch
72 conferred lysozyme sensitivity and increased swarming but did not rescue virulence defects.
73 O-antigen ligase, that resulted in a loss of swarming but not swimming motility.
74                                              Swarming (but not swimming) gene expression and motility
75 is toward nutrient has been thought to drive swarming, but here the nature of swarm growth and the im
76 Salmonella enterica, absence of FlhE affects swarming, but not swimming, motility.
77    Thus, TFP physically affect P. aeruginosa swarming by actively promoting cell-cell association and
78 fore, although O antigen may serve a role in swarming by promoting wettability, the loss of O antigen
79 red that elevated levels of c-di-GMP inhibit swarming by skewing stator selection in favor of the non
80 lution assays showed that repeated rounds of swarming by wildtype Pf-5 drives the accumulation of gac
81 gellar genes were highly upregulated in both swarming cells and consolidation-phase cells.
82                         Direct comparison of swarming cells to consolidation-phase cells found that 5
83                                              Swarming cells' speeds are comparable to bulk swimming s
84 lly expressed in broth-cultured cells versus swarming cells, and 527 genes were differentially expres
85               Fimbriae were downregulated in swarming cells, while genes involved in cell division an
86 l morphology with highly flexible snake-like swarming cells.
87  plethora of collective phenomena, including swarming, clustering, and phase separation.
88                                              Swarming colonies of independent Proteus mirabilis isola
89 GacS/GacA two-component regulatory system in swarming colonies of Pseudomonas protegens Pf-5.
90 n avenue for long-range communication in the swarming colony, ideally suited for secretory vesicles t
91 os of at least 2:1 lead to a collapse of the swarming colony.
92 erstating to the regulation required to make swarming cooperation stable.
93  on normal swarm agar in addition to being a swarming cue under normally nonpermissive conditions.
94 d, a plaP null mutation resulted in a modest swarming defect and slightly decreased levels of intrace
95   Higher flagellar numbers also suppress the swarming defect of mutants with changes in the chemotaxi
96 it pilA gene, show robust suppression of the swarming defect of the Delta bifA mutant, as well as its
97                        Here we show that the swarming defect of these mutants can also be suppressed
98 y, we isolated suppressors of the Delta bifA swarming defect.
99 e (efp) primarily supports Bacillus subtilis swarming differentiation, whereas EF-P in Gram-negative
100 lla propel bacteria during both swimming and swarming, dispersing them widely.
101  These findings broaden our understanding of swarming dynamics and have implications for the engineer
102                                Investigating swarming dynamics requires high-resolution imaging of si
103  which may explain their predominance on the swarming edge.
104 tion of gacS/gacA spontaneous mutants on the swarming edge.
105                                 The study of swarming elucidates more than proximate mechanisms: It e
106      In the animal kingdom, various forms of swarming enable groups of autonomous individuals to tran
107 s participates in two functions critical for swarming, enabling hydration and overriding surface fric
108 t that mixing between roosts during seasonal swarming events is necessary to maintain EBLV-2 in the p
109 pic effects in virulence phenotypes (reduced swarming, exo-protease and pyocyanin production).
110       As an example, we show that neutrophil swarming experiments exhibit dynamical signatures consis
111          The role of pseudodesmin as a major swarming factor was corroborated by identification and i
112 essed the signals that promote initiation of swarming following initial contact with a surface.
113                                Additionally, swarming from colonies grown on MacConkey soft agar was
114 d produce colonies with more deeply branched swarming fronts than the wild type or the sscL mutant.
115 e supernatants and demonstrated to stimulate swarming gene expression at low cell density.
116 s from exploratory patrolling to coordinated swarming, giving rise to dense clusters that further dis
117  genus Schistocerca, which contains both non-swarming grasshoppers and swarming locusts.
118 sts acquire key behavioral characters of the swarming gregarious phase within just 1 to 4 h of forced
119                                              Swarming has been implicated in pathogenesis; however, i
120                             Experiments with swarming have unveiled a strategy called metabolic prude
121 sensing, pattern formation, and emergence of swarming in active Caenorhabditis elegans aggregates.
122  algae B516, which inhibits V. alginolyticus swarming in its vicinity.
123 ative amino acid decarboxylase that inhibits swarming in Proteus mirabilis.
124 ion of cooperative rhamnolipids required for swarming in Pseudomonas aeruginosa.
125 ow that P. mirabilis CaUTI isolates initiate swarming in response to specific nutrients and environme
126    Testing of isogenic mutants revealed that swarming in response to the cues required putrescine bio
127           Suppressor mutations that enhanced swarming in the absence of YmfI were found at two positi
128 of the factors that contribute to neutrophil swarming in the extravascular space of a damaged tissue.
129 catone), octanal, nonanal and decanal during swarming in the laboratory.
130 ding arginine decarboxylase, is required for swarming in the urinary tract pathogen Proteus mirabilis
131                                              Swarming in the waaL mutant was restored by overexpressi
132 ted levels of c-di-GMP as well as stimulates swarming in the wild-type strain, while overexpression o
133  we report that quorum sensing can stimulate swarming in V. parahaemolyticus; it does so via an alter
134 is a differential effect of RcsF and UmoB on swarming in wild-type and waaL backgrounds, (ii) RcsF in
135  two morphological changes that occur during swarming--increases in cell length and flagellum density
136 eractions are critical for c-di-GMP-mediated swarming inhibition.
137 le, which was named avaroferrin, as a potent swarming inhibitor.
138 rric iron (Fe(3+)) availability to determine swarming initiation and biofilm formation.
139 +) to switch off RssAB signaling, triggering swarming initiation and biofilm reduction.
140 sAB signaling activation, leading to delayed swarming initiation and increased biofilm formation.
141 ntary form of self-organization occurs among swarming insects, flocking birds, or schooling fish; now
142                       From flocking birds to swarming insects, interactions of organisms large and sm
143                                              Swarming is a conspicuous behavioral trait observed in b
144                                              Swarming is a form of collective bacterial motion enable
145                                              Swarming is a phenomenon where collective motion arises
146                                 We show that swarming is abolished above a critical fraction of non-a
147                                   Precocious swarming is due to an increase in the number of elongate
148                 Our simulations suggest that swarming is simply driven by local food depletion but ot
149 growth that is distinct from the biofilm and swarming lifestyles of Salmonella.
150 dition to the well-characterized biofilm and swarming lifestyles, bacteria can also develop as micro-
151 on ancestor of Schistocerca must have been a swarming locust that crossed the Atlantic Ocean from Afr
152  contains both non-swarming grasshoppers and swarming locusts.
153 study, and the consequences of multispecies, swarming logistics networks.
154 rexpressing fcrX(+) was sufficient to induce swarming, luminescence and iron uptake gene expression i
155 flux highlights the important role of large, swarming macrozooplankton in POC export and, the need to
156 hythmic behaviors including flight activity, swarming, mating, host seeking, egg laying, and sugar fe
157                                          The swarming mechanism relies on the interaction between ind
158 ificantly upregulated in DeltagacA mutant on swarming medium.
159 and new data on honeybee colony development, swarming, mortality, pathogens and immune gene expressio
160                                 Swimming and swarming motilies in P. mirabilis were also significantl
161  been achieved in understanding swimming and swarming motilities powered by flagella, and twitching m
162 ctions - pilus biogenesis and multifactorial swarming motility - while shaping distinct nanoscale (bi
163 at is known about the proteins that modulate swarming motility and appear to act upstream of the mast
164                        In addition, in vitro swarming motility and quorum sensing signal production w
165 is requires only the MotA/MotB stator during swarming motility and that the residues required for sta
166 sing-controlled phenotypic traits, including swarming motility and the production of rhamnolipid and
167 for mutations in a single gene that improves swarming motility at the expense of biofilm formation.
168 ellar density above a critical threshold for swarming motility atop solid surfaces.
169 sites specifically favor flagellar motility, swarming motility based on 3-(3-hydroxyalkanoyloxy) alka
170 sensitivity to cysteine toxicity and altered swarming motility but unaltered cysteine-enhanced antibi
171                                              Swarming motility by the urinary tract pathogen Proteus
172 easuring extracellular protease activity and swarming motility confirmed the in vitro phosphorylation
173  detected by traditional methods yet enabled swarming motility in a strain that exhibited deficient p
174 tching motility, rhamnolipid production, and swarming motility in P. aeruginosa.
175 re, deletion of rgsA significantly increased swarming motility in P. aeruginosa.
176 et al. offer a new, previously unseen way of swarming motility inhibition in Pseudomonas aeruginosa P
177 nthesis SwrA governed by the adaptor protein swarming motility inhibitor A (SmiA).
178                                              Swarming motility is a flagella-driven multicellular beh
179            Movement over an agar surface via swarming motility is subject to formidable challenges no
180                                              Swarming motility is the movement of bacteria over a sol
181 lW or pilX gene alleviates the inhibition of swarming motility observed for strains with elevated lev
182 (2+) to the agar also partially restored the swarming motility of the znuC::Kan strain, but the addit
183 roduction, alginate production, swimming and swarming motility of uropathogens.
184       A mutation in yenI strongly stimulated swarming motility on the surface of semi-solid agar, whi
185 erium well known for its flagellum-dependent swarming motility over surfaces.
186 lar growth kinetics, whereas the microscopic swarming motility phases are dominated by physical cell-
187 e surface-induced (not repressed) and encode swarming motility proteins, virulence factors or sensory
188          Here we show that the repression of swarming motility requires a functional MotAB stator com
189 sed of EepR and EepS regulates hemolysis and swarming motility through transcriptional control of the
190            Mutating the motAB genes restores swarming motility to a strain with artificially elevated
191 ntaneous mutants were isolated that restored swarming motility to L. monocytogenes secA2 mutants.
192                                              Swarming motility was also restored by the addition of Z
193  in linking the partner-switching system and swarming motility was established by analyzing the swarm
194              The production of flagellin and swarming motility were restored by complementation with
195 isplayed enhanced collagenase activity, high swarming motility, and a destructive phenotype against c
196 es, have septation defects, are impaired for swarming motility, and form small plaques in tissue cult
197 lation, contact-dependent growth inhibition, swarming motility, and induction of antibiotic resistanc
198 nimal medium, displayed reduced swimming and swarming motility, and produced less flaA transcript and
199 uding resistance to host killing mechanisms, swarming motility, and protease production.
200 nges including delayed growth rate, retarded swarming motility, and pyocyanin overproduction.
201 luding biofilm formation, flagellum-mediated swarming motility, and type IV pilus-driven twitching.
202 ulates the same phenotypes as BDSF including swarming motility, biofilm formation, and virulence.
203 e data, a DeltadipA mutant exhibited reduced swarming motility, increased initial attachment, and pol
204  production, high collagenase activity, high swarming motility, low resistance to chloramphenicol, an
205 dition, ppGpp deficiency resulted in loss of swarming motility, reduction of pyoverdine production, i
206 ve histidine kinase), and PAO1 DeltaalgR for swarming motility, rhamnolipid production, and rhlA tran
207 rsely controls biofilm formation and surface swarming motility, with high levels of this dinucleotide
208  regulate genes for biofilm formation or for swarming motility-the output phenotypes.
209  has been shown to repress both swimming and swarming motility.
210 protein complexes necessary for swimming and swarming motility.
211 rexpression of MotA from a plasmid represses swarming motility.
212 PNPase mutant and also predicted a defect in swarming motility.
213 uired for the putrescine-dependent rescue of swarming motility.
214 ve over agar surfaces by flagellum-dependent swarming motility.
215 lus assembly machinery, in the repression of swarming motility.
216 lator activity have a commensurate effect on swarming motility.
217 oxidative stress, antibiotic resistance, and swarming motility.
218 -yet-unknown downstream mechanism to repress swarming motility.
219 rain had no effect on swimming but abolished swarming motility.
220 p behaviors, including biofilm formation and swarming motility.
221 system also contributes to the regulation of swarming motility.
222 hat YenS plays a direct, stimulatory role in swarming motility.
223  inverse regulation of biofilm formation and swarming motility.
224 ols mucoidy, while LadS negatively regulates swarming motility.
225 ss 3 promoters was found to be important for swarming motility.
226  aeruginosa, including biofilm formation and swarming motility.
227 ced biofilm formation but repressed swimming/swarming motility.
228 of 5-aminopentanonated EF-P is inhibitory to swarming motility.
229 uired for a form of surface migration called swarming motility.
230 32) of B. subtilis EF-P that is required for swarming motility.
231 olyl motifs present in proteins required for swarming motility.
232 ormation, quorum responses, development, and swarming motility.
233  as colony morphology, matrix formation, and swarming motility.
234 y of biological groups form a self-organized swarming motion at some point during their life spans, w
235                                              Swarming not only allows bacteria to forage for food, bu
236 ir migration in a process reminiscent of the swarming observed in neutrophils.
237 re systems in which both synchronization and swarming occur together.
238          We thus conclude that initiation of swarming occurs in response to specific cues and that ma
239      Antarctic krill (Euphausia superba) are swarming, oceanic crustaceans, up to two inches long, an
240 end composed of synthetic analogues to these swarming odours proved highly attractive to virgin males
241 cterial flagellum, powering the swimming and swarming of many motile bacteria.
242                              Because initial swarming of neutrophils at the site of infection occurs
243 llular role of TFP during flagellar-mediated swarming of P. aeruginosa that does not require TFP exte
244                                              Swarming on agar to which chloramphenicol had been added
245 ine was found to be a strict requirement for swarming on normal swarm agar in addition to being a swa
246 histidine, malate, and DL-ornithine promoted swarming on several types of media without enhancing swi
247 ore food; similarly, during large population swarming, only worms at the migrating front are in conta
248                      We find that neutrophil swarming over Candida clusters delays germination throug
249 ked for defects in swimming through liquids, swarming over surfaces, and transcriptional regulation.
250           Our simulations showed that TFP of swarming P. aeruginosa should be distributed all over th
251 mainly use computer simulations to study the swarming phenomena, this paper provides an analytical me
252 ed genes was altered consistent with the non-swarming phenotype observed for the mutant.
253 ng motility was established by analyzing the swarming phenotype of the PA3347 knock-out mutant and it
254 of the master regulators is important to the swarming phenotype perhaps at the level of controlling f
255 r exhibiting significant variation in social swarming phenotypes and five harboring large variation i
256                            Comparison of the swarming phenotypes of the single and double mutants and
257                                              Swarming populations exhibit non-genetic or adaptive res
258 eactions exhibit collective behavior such as swarming, predator-prey interactions, and chemotaxis tha
259 y, over 90% for each method, with successful swarming prediction up to 30 days prior to the event.
260 es and nutrient requirements involved in the swarming process have been identified, few studies have
261 hod for the monitoring and predicting of the swarming process within honeybee colonies, using vibro-a
262                 It is puzzling however, that swarming-proficient and virulent strains of Vibrio parah
263               In these studies, we examine a swarming-proficient, virulent strain and identify an alt
264 lar c-di-GMP leading to transcription of the swarming program.
265 es of nanomotors, which assemble in distinct swarming regions, is illustrated.
266                                              Swarming represents a special case of bacterial behavior
267                                 Furthermore, swarming repression by PilY1 specifically requires the d
268     We show that this HHQ- and PCA-dependent swarming repression is apparently independent of changes
269 ly but still functional in c-di-GMP-mediated swarming repression, indicating our ability to separate
270     This research investigates the effect of swarming season and sourcing geographical area on the nu
271 onal spectra predict the swarming within the swarming season only, and that this limitation can be li
272 he occurrence of queen pipes over the entire swarming season.
273                    Furthermore, we show that swarming species do not form a monophyletic group and no
274 ies do not form a monophyletic group and non-swarming species that are closely related to locusts oft
275 their composition and remain in a collective swarming state or even differentiate among behavioral ph
276                                              Swarming strategies are as diverse as the bacteria that
277 ageenan agar that may lay the foundation for swarming studies of snake-like, nonrod-shaped motile cel
278 ed (tan) colonies but still phase varied for swarming suggesting that pigmentation is not the cause o
279 rve the swirling that is conspicuous in many swarming systems, probably due to increasingly long-live
280 ned syringafactin production exhibited broad swarming tendrils, while a syringafactin-producing strai
281 rain that overproduced HAA exhibited slender swarming tendrils.
282 ways, forms branched tendril patterns during swarming; this phenomena occurs only when bacteria produ
283                           Avaroferrin blocks swarming through its ability to bind iron in a form that
284 es have suggested that FliL is essential for swarming through its involvement in viscosity-dependent
285 duction of the capsule and the repression of swarming to elucidate the global scope of genes in the O
286 teria move in groups, in a mode described as swarming, to colonize surfaces and form biofilms to surv
287 wer cell density) between antagonist strains swarming toward each other.
288 pt, including swimming in aqueous media, and swarming, twitching and gliding on solid and semi-solid
289 eduction correlates with increased NO-driven swarming, underlining a significant role for this motili
290 ype IV pilus biogenesis, function to repress swarming via modulation of intracellular c-di-GMP levels
291 age series to capture the motion dynamics of swarming Vibrio alginolyticus at cellular resolution ove
292 understand the molecular signals controlling swarming, we isolated two bacterial strains from the sam
293        Inspired by the dynamics of bacterial swarming, we report a swarm of polymer-brush-grafted, gl
294 y to flagella-dependent migration modes like swarming, we show that this much faster "colony surfing"
295 ates, were tested for the ability to promote swarming when added to normally nonpermissive media.
296 al suspensions, granular materials and crowd swarming, where consequences may be dramatic.
297             Here, we focus on two aspects of swarming, which, if understood, hold the promise of reve
298                    OBD programmed neutrophil swarming with an increase in isoprostanoid levels, with
299 nstantaneous vibrational spectra predict the swarming within the swarming season only, and that this
300 In this work, we show the iron regulation of swarming works through the ferric uptake regulator prote

 
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