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1 and within-group cooperation (e.g. parochial altruism).
2 at a cost to themselves, a phenomenon termed altruism.
3  sufficient for the evolution of one form of altruism.
4 onsumption of a common resource is a form of altruism.
5 r selfishness help to "pay" for second-order altruism.
6  than others, a situation called competitive altruism.
7 e punisher, making it a form of second-order altruism.
8 us action performance predicts self-reported altruism.
9 e punisher, making it a form of second-order altruism.
10 wo alleles which code for different kinds of altruism.
11 n alternative route towards the evolution of altruism.
12 m interspersed with phases of transient true altruism.
13 mpetition could account for the evolution of altruism.
14  are highly unstable, leading to the loss of altruism.
15 encouraging the origin and stability of true altruism.
16 the system towards the point of 50% marginal altruism.
17 y different perspectives on the evolution of altruism.
18 relationship between sexual reproduction and altruism.
19 relationship between sexual reproduction and altruism.
20 s and cultural context in the development of altruism.
21  are motivated more by self-interest than by altruism.
22 l infinite models) plays in the evolution of altruism.
23 lassic "haystack" models of the evolution of altruism.
24 ups contribute to research solely because of altruism.
25 n relatives can counteract kin selection for altruism.
26 d in terms of extended kinship or reciprocal altruism.
27 upport the selection of both weak and strong altruism.
28 ffects can cancel, limiting the viability of altruism.
29 tly responding quantitatively to a partner's altruism.
30 on need to be interpreted solely in terms of altruism.
31 table component of the immune system and kin altruism.
32 relatedness), group selection and reciprocal altruism.
33 es several novel insights into the nature of altruism.
34 s been described in terms of cooperation and altruism.
35 le of the neuropeptide oxytocin in promoting altruism.
36 sibility of a neural basis for extraordinary altruism.
37 eciprocal activity yielded little subsequent altruism.
38  is applied to model the real motivations of altruism.
39 ew that competition puts any such demands on altruism.
40  close kin but share a "greenbeard" gene for altruism?
41                             The evolution of altruism, a behaviour that benefits others at one's own
42 withstanding that examples of pathologies of altruism abound.
43 nces in the social preferences that motivate altruism across the primate order, and there is currentl
44 ection can favor reproductive altruism if an altruism allele aids copies of itself by helping relativ
45                     Concepts of pathological altruism, altruism bias, and guardian systems may help o
46 or kinship per se to achieve cooperation and altruism among group members.
47         Competition hinders the evolution of altruism amongst kin when beneficiaries gain at the expe
48                            With this type of altruism, an offspring will perform any act for which th
49  is unlikely that a single gene can code for altruism and a recognizable tag.
50 ing a common prosocial motivation underlying altruism and cooperation.
51 re, scientifically informed understanding of altruism and cooperative behavior.
52 istic scenarios, competition influences both altruism and defection.
53          Heritable variation in selfishness, altruism and discrimination is predicted to be particula
54 us social preferences in terms of both their altruism and equality-efficiency tradeoffs.
55 els of the gene-culture coevolution of human altruism and further sharpen what any theory of human co
56            Our study suggests that parochial altruism and in-group/out-group biases emerge early duri
57              Second, we ask why reproductive altruism and individuality arise only in the larger memb
58 e of cooperative agreements, from reciprocal altruism and insurance arrangements to the social norms
59 ormation, allowing the ethical principles of altruism and justice to guide organ allocation.
60  were thought to potentially benefit through altruism and maintenance of hope.
61 sness was inversely predictive of children's altruism and positively correlated with their punitive t
62           We show in a simulation model that altruism and punishment paradoxically become negatively
63 nstitutions of integration can unleash human altruism and restore cooperation in the presence of dive
64                                              Altruism and selfishness are 30-50% heritable in man in
65          This genetically based variation in altruism and selfishness requires explanation.
66 ions initially in stable equilibrium between altruism and selfishness.
67 eories on the explanation of both biological altruism and sex-ratio conflicts, and defend that the en
68 veral fundamental issues in the evolution of altruism and spite have remained contentious.
69 Humans are a cooperative species, capable of altruism and the creation of shared norms that ensure fa
70                            Here, we assessed altruism and third-party evaluation of scenarios depicti
71                                    Both pure altruism and warm-glow motives appear to determine the h
72 cussion of the connection between other-only altruism and whole-group altruism, in which the donor ga
73  relatedness predisposes individuals towards altruism, and as haploid germ cells of an ejaculate will
74 dual-level ones, need not produce behavioral altruism, and do not require competition between groups
75 as political mobilization, health practices, altruism, and emotional states exhibit similar dynamics
76  entered their children for medical benefit, altruism, and hope of cure.
77 cision on a choice between self-interest and altruism, and if improving this sensitivity through trai
78 ad private insurance, showed lower levels of altruism, and less understanding of study design.
79 enefit from medical improvement, feelings of altruism, and maintenance of hope, the chance of cure or
80  structure for the maintenance of mutualism, altruism, and niche construction or ecosystem engineerin
81 n ambivalent about their values (protection, altruism, and respect) and the deceased's wishes, not wa
82 m benefits the group, selfishness undermines altruism, and the purpose of the model is to identify me
83   METHODS/PRINCIPAL FINDINGS: In our AlAn's (altruism-antisocial) game a computer program presents su
84 eard effect and find that if recognition and altruism are always inherited together, the dynamics are
85 despite the fact that virtually all forms of altruism are associated with tradeoffs--some of enormous
86 ve been proposed to explain the evolution of altruism are direct reciprocity and indirect reciprocity
87  related individuals so that the benefits of altruism are reaped by copies of the altruistic gene in
88 cial interactions, including cooperation and altruism, are characteristic of numerous species, but ma
89 ong evolutionary trajectories, and find that altruism arises before directly beneficial behavior, des
90           Participants acknowledged hope and altruism as reasons for entering the trial more than exp
91 enomena, chiefly the evolution of biological altruism as that found in sterile castes of eusocial ins
92 netically based variation in selfishness and altruism, as in man, altruists with an innate ability to
93 t to the system's evolved functions: sibling altruism, aversion to personally engaging in sibling inc
94                                     And, can altruism be favored between individuals who are not clos
95  of competition, selection will often favour altruism because its alternatives provide lower fitness.
96     In the typical evolutionary formulation, altruism benefits the group, selfishness undermines altr
97 ed oxytocin system activity induces a social altruism bias at the cost of ecological responsibility.
98 eptual approach toward the quantification of altruism bias is presented.
99           Concepts of pathological altruism, altruism bias, and guardian systems may help open many n
100 despite fear), and adaptive social behavior (altruism, bonding, and teamwork) were found to be releva
101 ial insect colonies are pinnacles of evolved altruism but also exhibit dramatic conflict among relati
102 sity of the population slows up selection of altruism, but does not affect its direction, and this ho
103 mportant mechanism favoring the evolution of altruism, but punishment can be costly to the punisher,
104 uently relied on kin selection or reciprocal altruism, but recent models suggest that guarding may be
105 e hypothalamic peptide oxytocin in promoting altruism, but whether the influence of oxytocin benefits
106   This effect both supports the evolution of altruism by focusing the altruists' gifts on relatives o
107 provide a new perspective on strong vs. weak altruism by identifying their different underlying game
108 conditions associated with outgroup-directed altruism by showing that charitable social cues co-occur
109   In a brood from a single egg, reproductive altruism by soldiers reflects clone-level allocation to
110 uestions about the proximate source of human altruism by suggesting that prosocial behavior results,
111 on amongst defectors nevertheless undermines altruism, by facilitating invasion of unrelated benefici
112 en helped by the beneficiary, discriminating altruism can be resistant against invasion by defectors.
113                                   Reciprocal altruism can become established among selfish, unrelated
114 m, the model shows that both strong and weak altruism can evolve in periodically formed random groups
115                         The fact that strong altruism can increase when groups are periodically and r
116     We show how kin selection and reciprocal altruism can promote cooperation in diverse 2x2 matrix g
117 losses suffered in the first and in this way altruism can ratchet up to high levels.
118         Together, a little bit of reciprocal altruism can, however, greatly reduce the threshold at w
119 olving an absolute cost to altruists (strong altruism) cannot evolve when populations are structured
120 ow religion negatively influences children's altruism, challenging the view that religiosity facilita
121  Much of what we know about the evolution of altruism comes from animals.
122             The study of prosocial behavior--altruism, cooperation, trust, and the related moral emot
123                              Empathy-induced altruism derives its strength from the emotional stake i
124                                    A species altruism directed toward repair of human problems is cou
125 s the essential factors for the evolution of altruism directly in its parameters and integrates impor
126                               Moreover, when altruism does occur among other primates, it is typicall
127                                      Whereas altruism drives the evolution of human cooperation, ethn
128                        This species exhibits altruism during both asexual and sexual cycles of its li
129                   In some cases however, kin-altruism effects appear to be modest.
130                                   Reciprocal altruism emerges from iterated games where players have
131                              This nepotistic altruism evolves under natural selection only if the rat
132               The study of human and primate altruism faces an evolutionary anomaly: There is ample e
133  support decisions about trust, reciprocity, altruism, fairness, revenge, social punishment, social n
134 rlapping generations hinder the evolution of altruism, fecundity effects are more conducive to altrui
135 iprocal interactions are a potent trigger of altruism for young children, and that these interactions
136 ibly accounting for the distinctive forms of altruism found in our species.
137  of such functions when observing refusal of altruism from a genetic sister.
138 xperience fostered children's expectation of altruism from others.
139 l design allows us to rigorously distinguish altruism from preferences regarding equality-efficiency
140              Cooperation based on reciprocal altruism has evolved in only a small number of species,
141  Efforts to solve the evolutionary puzzle of altruism have a lengthy history, and recent years have s
142    However, the potential hurtful aspects of altruism have gone largely unrecognized in scientific in
143 as been used extensively to study reciprocal altruism, here we show that the n-player prisoner's dile
144 owever, in conditions that do not select for altruism, host bacteria promoting transfer are outcompet
145 ate mechanism to underlie so-called directed altruism, i.e., altruism in response to anothers's pain,
146             Selection can favor reproductive altruism if an altruism allele aids copies of itself by
147 indirect fitness options and helping is only altruism if it reduces the helper's direct fitness.
148 reciprocal activity elicited high degrees of altruism in 1- and 2-y-old children, whereas friendly bu
149 vity to inequality is strongly predictive of altruism in an independent task domain.
150 o far, and I apply the theory of competitive altruism in arguing how strategic investment in behaviou
151  further our understanding of the origins of altruism in humans by highlighting the importance of emo
152 eted theoretical results on the selection of altruism in inelastic viscous homogeneous populations, n
153 is may explain reported genetic variation in altruism in man.
154                     The profound benefits of altruism in modern society are self-evident.
155 selection theory predicts that the degree of altruism in queenless colonies should be reduced because
156  underlie so-called directed altruism, i.e., altruism in response to anothers's pain, need, or distre
157 tions meet the most stringent definitions of altruism in that they represent an intentional behavior
158 st defectors presents relative advantages to altruism in the simplest games between altruists and def
159 fications of pathological altruism, that is, altruism in which attempts to promote the welfare of oth
160 of Hamilton's rule in the case of other-only altruism in which the benefits are shared by other membe
161 d, providing field evidence for 'competitive altruism' in which helpful acts are used as a display to
162         Here we demonstrate the evolution of altruism, in the form of conditional reproductive restra
163  between other-only altruism and whole-group altruism, in which the donor gains some benefit from its
164 other prominent theories of the evolution of altruism: inclusive fitness and multilevel selection.
165 s us to combine kin selection and reciprocal altruism into a general matrix game model.
166                                              Altruism is consequently deemed to require stronger kin
167               The evolution of sociality and altruism is enigmatic because cooperators are constantly
168 is caused by loosely coupled separate genes, altruism is facilitated through beard chromodynamics in
169                        In non-human animals, altruism is generally directed towards relatives, and sa
170                         This is true even if altruism is initially rare, migration between groups all
171 enome sequencing reveals that this bacterial altruism is made possible by drug-resistance mutations u
172                                   In humans, altruism is motivated at least in part by empathy and co
173                        Our findings show how altruism is preserved from the disruptive effects of suc
174                                If reciprocal altruism is to provide an explanation for altruistic beh
175 e for charitable contributions, called "pure altruism," is satisfied by increases in the public good
176  In the absence of kin selection, reciprocal altruism may be an evolutionarily stable strategy but is
177                                              Altruism may be learned (behavioral evolution) in a way
178 ltruist, and also limits the extent to which altruism may emerge by exposing clusters of altruists to
179 riodically and randomly formed suggests that altruism may evolve more readily and in simpler organism
180 support prior suggestions that self-reported altruism may not reliably predict altruistic behaviour.
181                     However, a self-reported altruism measure previously linked to peer evaluations b
182 riments, interoceptive sensitivity predicted altruism measured through monetary generosity.
183 l of remittances can be explained by our kin-altruism model.
184 ndividuals favour kin to the extent that kin-altruism models predict?
185  that if punishing inequity is predictive of altruism more broadly, extraordinary altruists should pu
186 aggregate of scores from Ego-Resiliency, NEO Altruism, NEO Straightforwardness (positive predictors)
187 itness of the actor and also either benefit (altruism) or harm (spite) other individuals.
188                                              Altruism (over social space) corresponds to self-control
189  parents, who had higher levels of trust and altruism, perceived the potential for enhanced care, ref
190 ons suggest that enforcement of reproductive altruism (policing) in hymenopteran insect societies is
191                                              Altruism presents a challenge to evolutionary theory bec
192 ions, and hence, perhaps surprisingly, serve altruism rather than selfishness.
193  explanations and models for cooperation and altruism--reciprocity, kin and group selection, and puni
194    Although the neural mechanisms underlying altruism remain unknown, empathy and its component abili
195 teria to humans: Is the evolution of extreme altruism, represented by the sterile workers of social i
196               In order to be able to enforce altruism, reproductive cheaters need to be reliably iden
197                                   Greenbeard altruism resolves this paradox by allowing cooperators t
198 h as cooperation, giving, and other forms of altruism--result from covert attempts to avoid social in
199                                              Altruism should be the guiding motivation for all donati
200 used to model Herbert Simon's explanation of altruism, showing that altruistic norms can "hitchhike"
201 origin of the genetic basis for reproductive altruism (somatic cells specialized at vegetative functi
202 social conformity in general and competitive altruism specifically.
203  selective pressures leading to reproductive altruism stem from the increasing cost of reproduction w
204 for altruistic sterility is favored when the altruism sufficiently benefits relatives carrying the ge
205 al responsibility emphasize the relevance of altruism, suggesting that more altruistic individuals ar
206 ism, fecundity effects are more conducive to altruism than survival effects, and one demographic regi
207  and potential ramifications of pathological altruism, that is, altruism in which attempts to promote
208 ndependently inherited loci, one controlling altruism the other discrimination, or a one locus model
209 range of puzzling behaviors, such as extreme altruism, the use of ethical principles, and romantic lo
210 redictions from kin selection and reciprocal altruism theory.
211 ral network positively reinforces reciprocal altruism, thereby motivating subjects to resist the temp
212    After a brief exploration on the basis of altruism, this review will discuss the assessment, evalu
213 gn, physician involvement, personal benefit, altruism, time, and incentives.
214 the positive assortment necessary for strong altruism to evolve does not require these additional mec
215 it defectors against each other allow strong altruism to evolve even in populations with negligible k
216  such as kinship or reciprocity, that enable altruism to evolve.
217 that spermatozoa may display cooperation and altruism to gain an advantage when inter-male sperm comp
218                                  This allows altruism to persist even in weakly structured population
219 should consider kin selection and reciprocal altruism together rather than as alternatives, and they
220                     The cowbird's unexpected altruism toward host offspring simply promotes its selfi
221 totally negate, the kin-selected benefits of altruism toward relatives.
222 ophobia generally failed to exhibit enhanced altruism toward the outgroup.
223 not affect its direction, and this holds for altruism towards any individual, not just immediate neig
224                                 By directing altruism towards kin, D. purpureum should generally avoi
225 dividuals will show less aggression and more altruism towards relatives.
226 iduals should show less aggression, and more altruism, towards closer kin.
227                Additional questions assessed altruism, trust, value for research, and perceptions of
228                       The idea of reciprocal altruism usually involves direct reciprocity: repeated e
229                         Consistent with pure altruism, we find that even mandatory, tax-like transfer
230 aphic regime (so-called death-birth) permits altruism whereas another (so-called birth-death) does no
231 e reciprocal cues remain potent elicitors of altruism, whereas a fourth study with preschoolers showe
232 n and obtain conditions for the stability of altruism which differ from Hamilton's rule by simply rep
233  neural bases of this unique aspect of human altruism, which extends beyond interpersonal interaction
234 y phenomena of human interactions: authentic altruism, why people cooperate intuitively, one-shot coo
235 udy is best described as egoistically biased altruism, with important implications for our understand
236  group formation, the origin of reproductive altruism within the group, and the further specializatio
237 ves, while defectors receive the benefits of altruism without providing any help in return.

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