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1 great apes (chimpanzee, bonobo, gorilla, and orangutan).
2 1 human haplotype and that of chimpanzee and orangutan.
3 1 human haplotype and that of chimpanzee and orangutan.
4  as well as in one common chimpanzee and one orangutan.
5 rative sequence was determined from a female orangutan.
6 ion of human and great-ape lineages from the orangutan.
7 ropeans) and in one chimpanzee, gorilla, and orangutan.
8 o sequenced in one common chimpanzee and one orangutan.
9  relative of human, chimpanzee, gorilla, and orangutan.
10 ationship of human, chimpanzee, gorilla, and orangutan.
11  age and the quantitative abilities of adult orangutans.
12  underlie the variation at alpha-2 globin in orangutans.
13 th an additional fixed substitution found in orangutans.
14 cognitive difference between chimpanzees and orangutans.
15 bdivision was identified within the Sumatran orangutans.
16 ed from a chimpanzee, gibbons, gorillas, and orangutans.
17 els of 10 Bornean orangutans and 19 Sumatran orangutans.
18 st, the locus is monomorphic in macaques and orangutans.
19 ave serum IgAs of gorillas, chimpanzees, and orangutans.
20 diversity in humans and chimpanzees, but not orangutans.
21 n recent integrations within chimpanzees and orangutans.
22  species, including humans, chimpanzees, and orangutans.
23 nnaires used earlier to rate chimpanzees and orangutans.
24                                         Four orangutans (1 juvenile, 2 subadults, and 1 adult) and te
25               Samples from 10 chimpanzees, 3 orangutans, 1 gorilla, 1 rhesus macaque, 1 mangabey, and
26 hesus monkeys (12-lipoxygenating ALOX15) and orangutans (15-lipoxygenating ALOX15), express an ALOX15
27 Old World primates (now found in the Bornean orangutan) a pericentric inversion and centromere shift
28 ite divergence estimates inferred from human-orangutan alignments of neutrally evolving genomic seque
29                    By using human-chimpanzee-orangutan alignments, inferences can be made about both
30                Under the assumption that the orangutan and African apes diverged 13 million years ago
31                                              Orangutan and chimpanzee field studies suggest that like
32 pecific KIRs evolved first, being present in orangutan and functionally like their human counterparts
33 gnized sequences from the human and from the orangutan and gibbon revealed wide overlap of elements a
34                In other species, such as the orangutan and gibbon, FISH signals were only identified
35  the following species: chimpanzee, gorilla, orangutan and macaque.
36 ed, including two important primate species, orangutan and marmoset, bringing the total to 46 assembl
37         New species in the past year include orangutan and six additional low coverage mammalian geno
38 eles and haplotypes for panels of 10 Bornean orangutans and 19 Sumatran orangutans.
39  63 chimpanzees, 11 bonobos, 48 gorillas, 37 orangutans and 2 gibbons and observed undescribed variat
40 cultural lives of chimpanzees, gorillas, and orangutans and consider the ways in which this knowledge
41 glutamine microsatellites in chimpanzees and orangutans and found three nonsynonymous single nucleoti
42 e also show that the non-human HBV clades in orangutans and gibbons resulted from cross-species trans
43 s of DNA from humans, chimpanzees, gorillas, orangutans and macaques (HCGOM), Patterson et al. sugges
44 sues within and between humans, chimpanzees, orangutans and rhesus macaques.
45 plying horizontal transfer of information in orangutans and show that a number of social learning pro
46 egion in chimpanzees, bonobos, a gorilla, an orangutan, and a baboon.
47 ogous U2-LTRs in human, chimpanzee, gorilla, orangutan, and baboon and examined numerous homologs of
48 ne in common and pygmy chimpanzees, gorilla, orangutan, and baboon was sequenced to study the evoluti
49  of tandemly repeated sequence in the human, orangutan, and chimpanzee as well as a loss of approxima
50  a CMT1A-REP-like sequence, whereas gorilla, orangutan, and gibbon have a single copy.
51 ious nonhuman primates (chimpanzee, gorilla, orangutan, and gibbon).
52 king sequence in human, chimpanzee, gorilla, orangutan, and macaque genomes.
53 FISH analyses of human, chimpanzee, gorilla, orangutan, and macaque reveal qualitative and quantitati
54 terparts and to available sequenced gorilla, orangutan, and Old World monkey counterparts, and, on a
55 analyzed in one chimpanzee, one gorilla, one orangutan, and one Old World monkey.
56 o had potentially viable ORFs in chimpanzee, orangutan, and rhesus macaque, underscoring their potent
57 A array using probes from human, chimpanzee, orangutan, and rhesus.
58 uman, common and pygmy chimpanzees, gorilla, orangutan, and siamang) were analyzed.
59 human nuclear background, whereas mtDNA from orangutan, and species representative of Old-World monke
60  a biochemical effect of the substitution in orangutans, and because of its presence solely in the Su
61 ed methods in the tissues of rhesus monkeys, orangutans, and humans.
62 s no features suggesting clear affinities to orangutans, and instead reveals a morphological pattern
63  slow growth and low rate of reproduction in orangutans, and may be an evolutionary response to sever
64 rons of the AMGX and AMGY genes from humans, orangutans, and squirrel monkeys and estimated that the
65 ion of the human and rhesus and three in the orangutan, any one of which could be responsible for ina
66 nd Western gorillas and Sumatran and Bornean orangutans appear to have experienced gene flow since th
67                                              Orangutans are the largest habitually arboreal mammal.
68 their noncoding flanking sequences in human, orangutan, baboon, and colobus.
69 primate species (human, chimpanzee, gorilla, orangutan, baboon, rhesus, and macaque) in a way that is
70 everal primates: human, chimpanzee, gorilla, orangutan, baboon, rhesus, and macaque.
71  have generally agreed it should be defined, orangutans behaved as individual and creative problem so
72  we document similar geographic variation in orangutan behaviors.
73 anzee individual, with data for gorillas and orangutans being anecdotal.
74 factor of activated T cells); chimpanzee and orangutan BILF1 molecules were unable to activate NFAT.
75 , 17, and 17 repeats in the gibbon, gorilla, orangutan, bonobo, neanderthal, and human Liat1, respect
76 al gyrus of chimpanzee, bonobo, gorilla, and orangutan brains through direct cytoarchitectonic compar
77                         Results suggest that orangutans but not squirrel monkeys possess Stage 6 obje
78 green pigment genes of Old World monkeys and orangutans but was present in intron 1 of both the green
79 hat of chimpanzees and bonobos, gorillas and orangutans by approximately 400, 635 and 820 kcal day(-1
80                                          The orangutan C1-specific KIR reacts or cross-reacts with al
81   The results of this study demonstrate that orangutans can recognize, or learn to recognize, relevan
82                                          The orangutan chose 1 of the quantities, which was removed,
83 scovered that the well-known polymorphism of orangutan chromosome 12 was due to the presence of an EN
84                           Haplotypes from 28 orangutan chromosomes were collected from a 1.46-kilobas
85 ominids (chimpanzees, bonobos, gorillas, and orangutans), comparative studies suggest a surprising nu
86 quences from cat, lynx, elephant, gopher and orangutan complement the previous database of sequences
87 ic inversion is needed to derive the Bornean orangutan condition.
88  diversity in fossil and extant apes and for orangutan conservation and reintroduction programs.
89 on that may be interpreted in the context of orangutan conservation efforts.
90    We find no evidence for nestedness in the orangutan cultural data.
91                         Moreover, two of the orangutans demonstrated this within the first test trial
92 ive organization strategy was found, but the orangutans developed a right-to-left spatial response st
93                        Results indicate that orangutans did not display conservation in the strict se
94                                     Although orangutans did not show conservation in the strict sense
95                                              Orangutans did not use the simple strategy of selecting
96 onvergent adaptation between chimpanzees and orangutans (digital elongation) and comparatively little
97  internalized, BILF1 from the chimpanzee and orangutan displayed an altered cellular localization pat
98            Our results suggest that although orangutans do make use of disturbed forest, they select
99 e currently recognized: Sumatran and Bornean orangutans, eastern and western gorillas, and chimpanzee
100                  Here, we show that Sumatran orangutans employ unique locomotor strategies to control
101                   These results suggest that orangutans exhibit a degree of technical knowledge and c
102 related "great apes" (bonobos, gorillas, and orangutans) express several CD33-related Siglecs on thei
103 ere, we test this hypothesis comparing human/orangutan genome-wide non-coding divergence (K) to that
104                           Our analyses of 37 orangutan genomes provided a second line of evidence.
105 lite births/deaths in human, chimpanzee, and orangutan genomes, using macaque and marmoset as outgrou
106                This is particularly true for orangutans (genus: Pongo), the only Asian great apes and
107 NA and mtDNA encoded complex I subunits from orangutan, gorilla, chimpanzee, human and all available
108 duced vocalizations from infant and juvenile orangutans, gorillas, chimpanzees, and bonobos, as well
109 and species-associated chimpanzee and gibbon/orangutan groups.
110 ound to carry three or four GAA repeats, the orangutans had four or five GAA repeats, and the gorilla
111                         In this context, the orangutan has particular relevance because it represents
112                                Although each orangutan has spontaneously used tools in the past, the
113                We sequenced the genome of an orangutan homozygous for the ENC, and we focused our ana
114  in 16 chimpanzees, 3 bonobos, and 2 Bornean orangutans; however, 9 chimpanzees and 6 Sumatran orangu
115 e distantly related species, including human-orangutan, human-baboon, and human-macaque.
116 rved differences in energy expenditure among orangutans, humans, and other mammals reflect known diff
117 molecular evolution compared to gorillas and orangutans in the regions analyzed.
118           Despite activity levels similar to orangutans in the wild, Great Ape Trust orangutans used
119 mpanzees, four bonobos, 14 gorillas, and six orangutans, in which interpretable MSY sequence ranges f
120                                              Orangutan inhibitory MHC-C-reactive KIRs pair with activ
121 ned analyses support a new classification of orangutans into three extant species.
122 n which chimpanzee is closest to the humans; orangutan is a clear outgroup of human, chimpanzee, and
123                                          The orangutan is the world's largest arboreal mammal, and im
124 ctive well-being in humans, chimpanzees, and orangutans is ancestral in catarrhine primates.
125 ranio-mandibular and dental characters of an orangutan killed in a human-animal conflict to those of
126                                              Orangutan KIR corresponding to the three KIR lineages ex
127                                          The orangutan KIR haplotypes have between 5 and 10 KIR genes
128                                              Orangutans lack C2 and C2-specific KIRs, but have a uniq
129                                    The seven orangutan lineage III KIR genes all locate to the centro
130 easured daily energy expenditure (kCal/d) in orangutans living in a large indoor/outdoor habitat at t
131 ors from EBV and LCVs from NHPs (chimpanzee, orangutan, marmoset, and siamang) were selected for mult
132            Results suggest that enculturated orangutans may not possess social-cognitive abilities si
133 ating isoforms are found in higher primates (orangutans, men), and these results suggest an evolution
134                                    For those orangutan MHC class I allotypes that are detected by hum
135               The data demonstrate that each orangutan MHC haplotype has one copy of the MHC-A gene,
136 e and complicated transcription profiles for orangutan MHC-A, -B, and -C.
137 his correlates with the observation that all orangutan MHC-C allotypes examined have the C1 motif.
138 movement among all age and sex classes, with orangutans more likely to move in directions of increase
139  of the flexibility of the supports on which orangutans moved.
140 rest canopy were the dominant determinant of orangutan movement among all age and sex classes, with o
141  (Light Detection and Ranging) to understand orangutan movement in disturbed and fragmented forests o
142  at least four occasions; in chimpanzees and orangutans, MSY2 contains only two units.
143                                Nest-building orangutans must daily build safe and comfortable nest st
144 Experiment 3, we endowed bonobos (N = 4) and orangutans (N = 5) with either one or 12 food items.
145 te the mechanical design and architecture of orangutan nests and determine the degree of technical so
146 onstrate that, compared to the average human/orangutan non-coding divergence (K=3%), the substitution
147                     On the other hand, human/orangutan non-coding divergence at the Xp/Yp pseudoautos
148  split in the evolutionary history of extant orangutans occurred approximately 3.38 mya between the B
149 an-animal conflict to those of 33 adult male orangutans of a similar developmental stage, we found co
150 ing humans, great apes (chimpanzee, gorilla, orangutan), Old- and New-World monkeys (macaque and marm
151 imary visual area in humans, chimpanzees, an orangutan, Old World monkeys, and New World monkeys.
152              In Experiment 2, performance of orangutans on double invisible displacements and control
153 zees previously tested on the same task, the orangutans optimized their performance.
154  proposed models ranging from African ape to orangutan or generalized Miocene ape-like.
155 es are structurally more derived than either orangutan or human genomes.
156 lete sequence of the chimpanzee, gorilla and orangutan orthologues should be deducible solely through
157          Designation of Bornean and Sumatran orangutans, P. pygmaeus (Linnaeus 1760) and P. abelii (L
158 ler cell Ig-like receptor (KIR) expressed by orangutan peripheral blood cells were examined by clonin
159                                          The orangutan Plasmodium parasite is a plausible selective a
160 data to evaluate the extent to which Bornean orangutans Pongo pygmaeus come down from the trees to tr
161 ts, protect adequate habitat for the Bornean orangutan (Pongo pygmaeus) and Bornean elephant (Elephas
162 CLC beta-galactoside binding sites from both orangutan (Pongo pygmaeus) and murine (Mus musculus) gen
163 y data for the critically endangered Bornean orangutan (Pongo pygmaeus), allowing a deeper understand
164 periments, chimpanzees (Pan troglodytes), an orangutan (Pongo pygmaeus), and human infants (Homo sapi
165  troglodytes), gorilla (Gorilla gorilla) and orangutan (Pongo pygmaeus).
166 roteins from EDN from sequences derived from orangutan (Pongo pygmaeus, oEDN) and Old World monkey (M
167 es, PTR), gorilla (Gorilla gorilla, GGO) and orangutan (Pongo pygmaeus, PPY).
168 al transmission of information in 15 captive orangutans (Pongo abelii and Pongo pygmaeus) using a sim
169 his study, chimpanzees (Pan troglodytes) and orangutans (Pongo abelii) either had to determine the lo
170 ist learning and memory to determine whether orangutans (Pongo pygmaeus x P. abelii) would show evide
171                           The authors tested orangutans (Pongo pygmaeus) and squirrel monkeys (Saimir
172 ve gorillas (Gorilla gorilla) and 19 captive orangutans (Pongo pygmaeus) and were compared with chimp
173       In this study, the authors presented 2 orangutans (Pongo pygmaeus) with a quantity judgment tas
174 from 2 gorillas (Gorilla gorilla gorilla), 4 orangutans (Pongo pygmaeus), 14 chimpanzees (Pan troglod
175 anzees (Pan troglodytes), a group of 2 adult orangutans (Pongo pygmaeus), and a group of 36 children
176 ences were studied for the two subspecies of orangutans (Pongo pygmaeus), which are located in Borneo
177  assessed in two human-reared (enculturated) orangutans (Pongo pygmaeus).
178 children, chimpanzees (Pan troglodytes), and orangutans (Pongo pygmaeus).
179 re we report daily energy use in free-living orangutans (Pongo spp.) and test whether observed differ
180 evelopment on biodiversity [1, 4, 5], and on orangutans (Pongo spp.) in particular, have been well do
181 rigid or flexible properties was explored in orangutans (Pongo spp.) through an extension of D.J. Pov
182 report reliable ages at M1 emergence for the orangutan, Pongo pygmaeus (4.6 y), and the gorilla, Gori
183 ly considered to be a relative of the extant orangutan, Pongo pygmaeus.
184           Our analysis revealed that Bornean orangutan populations have declined at a rate of 25% ove
185 important implications for the management of orangutan populations in captivity and in the wild, and
186 al value for the long-term sustainability of orangutan populations.
187 r putative cultural traits in chimpanzee and orangutan populations.
188 n bonobos, Western chimpanzees, and Sumatran orangutans-populations that have experienced recent gene
189       Orthologous to Patr-AL are polymorphic orangutan Popy-A and the 5' part of human pseudogene HLA
190 e is similar to the expressed A locus in the orangutan, Popy-A, suggesting they are orthologous.
191 Ala) and 15-lipoxygenating (man, chimpanzee, orangutan, rabbit, ratLeu353Phe) ALOX15 variants and fou
192  mouse, and adds seven additional organisms: orangutan, rat, cow, pig, horse, platypus and Arabidopsi
193                                              Orangutans react to branch flexibility like humans runni
194 proximately 9 Mb of human chromosome 21 with orangutan, rhesus macaque, and woolly monkey DNA sequenc
195  evolution of CD80 genes derived from human, orangutan, rhesus monkey, baboon, cat, cow, and rabbit b
196 dy, the molecular population genetics of the orangutan's alpha-2 globin (HBA2) gene were investigated
197 errestrial locomotion is part of the Bornean orangutan's natural behavioural repertoire to a much gre
198 provide additional information regarding the orangutan's position on the evolutionary tree of Pongida
199                                              Orangutans select different tactics for repairing failed
200 riant sites among the human, chimpanzee, and orangutan sequences.
201 utans; however, 9 chimpanzees and 6 Sumatran orangutans showed neither the 9.1-kb+ nor the 9.1-kb- al
202 howing population-level right-handedness and orangutans showing population-level left-handedness.
203                             In Experiment 1, orangutans solved all visible displacements and most inv
204  southernmost range limit of extant Sumatran orangutans south of Lake Toba, is distinct from other no
205 f the MHC class I gene repertoire in the two orangutan species, Pongo abelii and Pongo pygmaeus, is p
206 cause of its presence solely in the Sumatran orangutan species, the mutation may be associated with r
207 , found in both the Bornean and the Sumatran orangutan subspecies, was associated with different alte
208 ologous portions of CAPN10 in chimpanzee and orangutan (the identity of sites 43 and 19 was further i
209  C1-bearing MHC-C from MHC-B, as informed by orangutan, the focus changes to MHC-C and its cognate li
210 ficantly constrained relative to that of the orangutan, the Old World monkey, and the mouse, but not
211               In contrast, MHC haplotypes of orangutans, the Asian great ape species, exhibit variati
212                                  In Sumatran orangutans, the overall genomic diversity is approximate
213                                              Orangutans, therefore, attend to differences in magnitud
214  genes in 10 humans, one chimpanzee, and one orangutan to (i) provide an average estimate of nucleoti
215 15 million years in the higher primates from orangutan to human.
216                              The capacity of orangutans to come down from the trees may increase thei
217 n humans, chimpanzees, bonobos, gorillas and orangutans to test the hypothesis that the human lineage
218 t it allows the most arboreal great ape, the orangutan, to access supports too flexible to be negotia
219                               Chimpanzee and orangutan TRIM5alpha proteins functionally resembled hum
220 of these homologous sites was identified; in orangutan two sites were identified, while gibbon exhibi
221                          During construction orangutans use the fact that branches only break half-wa
222 r to orangutans in the wild, Great Ape Trust orangutans used less energy, relative to body mass, than
223 d cells purified from human, chimpanzee, and orangutan, using digestion with a methylation-sensitive
224 ific alpha satellite in chimpanzee, gorilla, orangutan, vervet, macaque, and baboon.
225 ition, the mutational spectrum in humans and orangutans was estimated as 63% AG (and CT), 17% AC (and
226               In contrast, chimpanzees and 1 orangutan were successful in using video to guide their
227                                        Older orangutans were less accurate than the young and middle-
228                                Three captive orangutans were presented with three unfamiliar pairs of
229 d canopy gap size influenced terrestriality, orangutans were recorded on the ground as frequently in
230 e in gene expression among human, chimp, and orangutan, which suggested an accelerated rate of diverg
231 two samples of chimpanzees and one sample of orangutans) whose well-being was assessed by raters fami
232 e in chimpanzees, one in gorillas and one in orangutans with derived allele frequencies of 0.01, 0.26
233 ghly detailed visual observations of Bornean orangutans with high-resolution airborne remote sensing
234  members (humans, chimpanzees, gorillas, and orangutans), with the most notable differences between t
235 ASMT (K=6.5%) genes located in the human and orangutan Xp/Yp pseudoautosomal region (p-PAR), where re

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