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1 liL cells also exhibit temperature-sensitive swarming.
2 t a particular environment is permissive for swarming.
3 agella as they colonize agar surfaces during swarming.
4 d, a means for surface colonization known as swarming.
5 ation of environmental signals that regulate swarming.
6 ability to inducing activity with respect to swarming.
7 his is a means for colony expansion known as swarming.
8 d that protein synthesis is not required for swarming.
9 h P. mirabilis prepares for the next wave of swarming.
10 ivity under selection for growth achieved by swarming.
11  solid surfaces by a type of motility called swarming.
12 ehavior, with ramifications beyond bacterial swarming.
13 low viscosity that normally impede wild-type swarming.
14 stimulating biofilm formation and repressing swarming.
15 rs, only one of which can provide torque for swarming.
16 e this question using Pseudomonas aeruginosa swarming, a collective surface motility requiring massiv
17 scriminate kin from nonkin in the context of swarming, a cooperative multicellular behavior.
18 ferent drying conditions along with the anti-swarming activity against Citrobacter rodentium.
19              Subspecies IIIa is nonmotile on swarming agar and thus may also have reduced motility un
20                                 Natural anti-swarming agents from food waste may have promising poten
21  upregulate flagellar gene expression during swarming, also do not increase flagellar numbers per mum
22 movement, confers an added advantage, making swarming an effective strategy for prevailing against an
23 ach colony is to produce all drones prior to swarming, an impossible solution on a population scale b
24 Deletion of MXAN_4832 causes defects in both swarming and aggregation related to cell motility and th
25 e upregulation of VLA3 to support neutrophil swarming and aggregation.
26                    Iron availability affects swarming and biofilm formation in various bacterial spec
27 ever, how bacteria sense iron and coordinate swarming and biofilm formation remains unclear.
28 mately play a role in surface-sensing during swarming and biofilm formation.
29 ias, which likely accounts for their reduced-swarming and delayed-development phenotypes.
30 developmental processes involving vegetative swarming and fruiting body formation.
31  the point mutants on single cell reversals, swarming and fruiting body formation.
32 uency to support directional motility during swarming and fruiting body formation.
33 sponse characterized by meningeal neutrophil swarming and microglial reconstitution of the damaged gl
34 he cellular mechanisms underlying neutrophil swarming and suggest new roles for neutrophils in shapin
35                                              Swarming and swimming motility of bacteria has been stud
36                                              Swarming and T3SS1 gene expression were also demonstrate
37 med CalR was identified and shown to repress swarming and T3SS1 gene expression.
38 rsals of gliding direction are essential for swarming and that reversals increase the outflow of cell
39 the physical mechanisms underlying bacterial swarming and the balance between individual and collecti
40 y, we used a computational model to simulate swarming and to probe for individual cell behavior that
41 laP and decreased both putrescine stimulated swarming and urothelial cell invasion in a speA mutant.
42  in Vibrio parahaemolyticus, in part because swarming and virulence factors--the hallmarks of the org
43   In contrast, tan variants are deficient in swarming (+) and persist beyond stationary phase.
44            Yellow variants are proficient at swarming (++) and tend to lyse in liquid during stationa
45           The bioactivity (antioxidant, anti-swarming) and phenolics content was significantly higher
46  by LafK, a sigma(54)-dependent regulator of swarming, and additionally connected by a negative-feedb
47       RNA was extracted from broth-cultured, swarming, and consolidation-phase cells to assess transc
48 is study provides detailed insights into the swarming architecture and dynamics of Vibrio alginolytic
49                                     These co-swarming assays further demonstrated that DeltagacA muta
50 tion of motility as measured in swimming and swarming assays.
51           Importantly, choline also enhances swarming-associated colony expansion of P. mirabilis und
52 oss of fliL also results in an inhibition of swarming at <30 degrees C.
53                                   We studied swarming at the cellular level using a combination of la
54 find nutrients and avoid toxic environments, swarming bacteria appear to suppress chemotaxis and to u
55                                              Swarming bacteria can be further subdivided by their req
56                                              Swarming bacteria move in multicellular groups and exhib
57                                              Swarming bacteria move on agar surfaces in groups, using
58                                              Swarming bacteria use kin discrimination to preferential
59 f the master flagellar regulators in diverse swarming bacteria.
60 e biosurfactants constitutively we show that swarming becomes cheatable: a non-producing strain rapid
61 ly applicable method for analyzing bacterial swarming behavior in two and three dimensions with both
62 nonmotile cells were sufficient to block the swarming behavior of a large gliding-proficient populati
63 energy-harvesting, environmentally triggered swarming behavior, and magnetic control of the new Janus
64 xis and cluster formation reminiscent of the swarming behaviour of insects.
65 duces coordinated, self-organized neutrophil-swarming behaviour that isolates the wound or infectious
66 ems (alternative sigma factors, sporulation, swarming, biofilm formation, stochastic cell fate switch
67 conferred lysozyme sensitivity and increased swarming but did not rescue virulence defects.
68 O-antigen ligase, that resulted in a loss of swarming but not swimming motility.
69                                              Swarming (but not swimming) gene expression and motility
70 is toward nutrient has been thought to drive swarming, but here the nature of swarm growth and the im
71 Salmonella enterica, absence of FlhE affects swarming, but not swimming, motility.
72    Thus, TFP physically affect P. aeruginosa swarming by actively promoting cell-cell association and
73 fore, although O antigen may serve a role in swarming by promoting wettability, the loss of O antigen
74 red that elevated levels of c-di-GMP inhibit swarming by skewing stator selection in favor of the non
75 lution assays showed that repeated rounds of swarming by wildtype Pf-5 drives the accumulation of gac
76 gellar genes were highly upregulated in both swarming cells and consolidation-phase cells.
77 (diffusing only a few micrometers) while the swarming cells streamed past underneath.
78                         Direct comparison of swarming cells to consolidation-phase cells found that 5
79                                              Swarming cells' speeds are comparable to bulk swimming s
80 lly expressed in broth-cultured cells versus swarming cells, and 527 genes were differentially expres
81               Fimbriae were downregulated in swarming cells, while genes involved in cell division an
82 l morphology with highly flexible snake-like swarming cells.
83                      Boundaries form between swarming colonies of different P. mirabilis strains but
84                                              Swarming colonies of independent Proteus mirabilis isola
85 GacS/GacA two-component regulatory system in swarming colonies of Pseudomonas protegens Pf-5.
86 n avenue for long-range communication in the swarming colony, ideally suited for secretory vesicles t
87 os of at least 2:1 lead to a collapse of the swarming colony.
88 erstating to the regulation required to make swarming cooperation stable.
89  on normal swarm agar in addition to being a swarming cue under normally nonpermissive conditions.
90                  At various times during the swarming cycle, the increased expression of wosA resulte
91 d, a plaP null mutation resulted in a modest swarming defect and slightly decreased levels of intrace
92   Higher flagellar numbers also suppress the swarming defect of mutants with changes in the chemotaxi
93 it pilA gene, show robust suppression of the swarming defect of the Delta bifA mutant, as well as its
94                        Here we show that the swarming defect of these mutants can also be suppressed
95 y, we isolated suppressors of the Delta bifA swarming defect.
96 e (efp) primarily supports Bacillus subtilis swarming differentiation, whereas EF-P in Gram-negative
97 lla propel bacteria during both swimming and swarming, dispersing them widely.
98 at although these genes play a minor role in swarming, dppA and cysJ are required during ascending ur
99  These findings broaden our understanding of swarming dynamics and have implications for the engineer
100                                Investigating swarming dynamics requires high-resolution imaging of si
101  which may explain their predominance on the swarming edge.
102 tion of gacS/gacA spontaneous mutants on the swarming edge.
103 s participates in two functions critical for swarming, enabling hydration and overriding surface fric
104                    We examined the motion of swarming Escherichia coli, comparing the motion of indiv
105 pic effects in virulence phenotypes (reduced swarming, exo-protease and pyocyanin production).
106 pattern formed by consecutive waves of rapid swarming followed by consolidation into shorter cells.
107 essed the signals that promote initiation of swarming following initial contact with a surface.
108 e supernatants and demonstrated to stimulate swarming gene expression at low cell density.
109  genus Schistocerca, which contains both non-swarming grasshoppers and swarming locusts.
110 tle-seen solitarious phase and the notorious swarming gregarious phase depending on population densit
111 sts acquire key behavioral characters of the swarming gregarious phase within just 1 to 4 h of forced
112                                              Swarming has been implicated in pathogenesis; however, i
113 red for swimming but completely defective in swarming in both organisms, and have studied this phenot
114  algae B516, which inhibits V. alginolyticus swarming in its vicinity.
115 enes, previously reported to be required for swarming in laboratory strains, are dispensable for robu
116 ative amino acid decarboxylase that inhibits swarming in Proteus mirabilis.
117 ion of cooperative rhamnolipids required for swarming in Pseudomonas aeruginosa.
118 ow that P. mirabilis CaUTI isolates initiate swarming in response to specific nutrients and environme
119    Testing of isogenic mutants revealed that swarming in response to the cues required putrescine bio
120           Suppressor mutations that enhanced swarming in the absence of YmfI were found at two positi
121 of the factors that contribute to neutrophil swarming in the extravascular space of a damaged tissue.
122 ding arginine decarboxylase, is required for swarming in the urinary tract pathogen Proteus mirabilis
123                                              Swarming in the waaL mutant was restored by overexpressi
124 ted levels of c-di-GMP as well as stimulates swarming in the wild-type strain, while overexpression o
125  we report that quorum sensing can stimulate swarming in V. parahaemolyticus; it does so via an alter
126 is a differential effect of RcsF and UmoB on swarming in wild-type and waaL backgrounds, (ii) RcsF in
127  two morphological changes that occur during swarming--increases in cell length and flagellum density
128 eractions are critical for c-di-GMP-mediated swarming inhibition.
129 le, which was named avaroferrin, as a potent swarming inhibitor.
130 rric iron (Fe(3+)) availability to determine swarming initiation and biofilm formation.
131 +) to switch off RssAB signaling, triggering swarming initiation and biofilm reduction.
132 sAB signaling activation, leading to delayed swarming initiation and increased biofilm formation.
133 ntary form of self-organization occurs among swarming insects, flocking birds, or schooling fish; now
134                       From flocking birds to swarming insects, interactions of organisms large and sm
135                                              Swarming is a conspicuous behavioral trait observed in b
136                                              Swarming is a phenomenon where collective motion arises
137                                 We show that swarming is abolished above a critical fraction of non-a
138                                   Precocious swarming is due to an increase in the number of elongate
139                           We first show that swarming is immune to the evolution of rhlA(-) 'cheaters
140 growth that is distinct from the biofilm and swarming lifestyles of Salmonella.
141 dition to the well-characterized biofilm and swarming lifestyles, bacteria can also develop as micro-
142 on ancestor of Schistocerca must have been a swarming locust that crossed the Atlantic Ocean from Afr
143  contains both non-swarming grasshoppers and swarming locusts.
144 study, and the consequences of multispecies, swarming logistics networks.
145 hythmic behaviors including flight activity, swarming, mating, host seeking, egg laying, and sugar fe
146                                          The swarming mechanism relies on the interaction between ind
147 ificantly upregulated in DeltagacA mutant on swarming medium.
148 tal conditions that may control the onset of swarming migration.
149                                 Swimming and swarming motilies in P. mirabilis were also significantl
150  been achieved in understanding swimming and swarming motilities powered by flagella, and twitching m
151 ctions - pilus biogenesis and multifactorial swarming motility - while shaping distinct nanoscale (bi
152  mM OPP, genes involved in the exhibition of swarming motility and anaerobic respiration were upregul
153 at is known about the proteins that modulate swarming motility and appear to act upstream of the mast
154 otropic defects in homologous recombination, swarming motility and cell division.
155                        In addition, in vitro swarming motility and quorum sensing signal production w
156                                         Weak swarming motility and rare flagella were observed in a m
157 is requires only the MotA/MotB stator during swarming motility and that the residues required for sta
158 sing-controlled phenotypic traits, including swarming motility and the production of rhamnolipid and
159 for mutations in a single gene that improves swarming motility at the expense of biofilm formation.
160 ellar density above a critical threshold for swarming motility atop solid surfaces.
161 sites specifically favor flagellar motility, swarming motility based on 3-(3-hydroxyalkanoyloxy) alka
162 sensitivity to cysteine toxicity and altered swarming motility but unaltered cysteine-enhanced antibi
163                                              Swarming motility by the urinary tract pathogen Proteus
164 easuring extracellular protease activity and swarming motility confirmed the in vitro phosphorylation
165  detected by traditional methods yet enabled swarming motility in a strain that exhibited deficient p
166 boratory strains, are dispensable for robust swarming motility in an undomesticated strain.
167  We redemonstrate that SwrA is essential for swarming motility in Bacillus subtilis, and we reassert
168 tching motility, rhamnolipid production, and swarming motility in P. aeruginosa.
169 re, deletion of rgsA significantly increased swarming motility in P. aeruginosa.
170 et al. offer a new, previously unseen way of swarming motility inhibition in Pseudomonas aeruginosa P
171 nthesis SwrA governed by the adaptor protein swarming motility inhibitor A (SmiA).
172  or a gene encoding a previously unannotated swarming motility inhibitor, SmiA.
173                                              Swarming motility is a flagella-driven multicellular beh
174            Movement over an agar surface via swarming motility is subject to formidable challenges no
175                                              Swarming motility is the movement of bacteria over a sol
176 lW or pilX gene alleviates the inhibition of swarming motility observed for strains with elevated lev
177 (2+) to the agar also partially restored the swarming motility of the znuC::Kan strain, but the addit
178 roduction, alginate production, swimming and swarming motility of uropathogens.
179 aaL gene, encoding O-antigen ligase, blocked swarming motility on solid surfaces but had little effec
180       A mutation in yenI strongly stimulated swarming motility on the surface of semi-solid agar, whi
181 erium well known for its flagellum-dependent swarming motility over surfaces.
182 e surface-induced (not repressed) and encode swarming motility proteins, virulence factors or sensory
183          Here we show that the repression of swarming motility requires a functional MotAB stator com
184 sed of EepR and EepS regulates hemolysis and swarming motility through transcriptional control of the
185            Mutating the motAB genes restores swarming motility to a strain with artificially elevated
186 ntaneous mutants were isolated that restored swarming motility to L. monocytogenes secA2 mutants.
187                                              Swarming motility was also restored by the addition of Z
188  in linking the partner-switching system and swarming motility was established by analyzing the swarm
189 essed motility; several distinct patterns of swarming motility were noted.
190              The production of flagellin and swarming motility were restored by complementation with
191  the exhibition of anaerobic respiration and swarming motility were upregulated.
192 s gene identified by its ability to increase swarming motility when overexpressed.
193 isplayed enhanced collagenase activity, high swarming motility, and a destructive phenotype against c
194 es, have septation defects, are impaired for swarming motility, and form small plaques in tissue cult
195 lation, contact-dependent growth inhibition, swarming motility, and induction of antibiotic resistanc
196 at functions in hydrophobic nutrient uptake, swarming motility, and pathogenesis.
197 nimal medium, displayed reduced swimming and swarming motility, and produced less flaA transcript and
198 uding resistance to host killing mechanisms, swarming motility, and protease production.
199 nges including delayed growth rate, retarded swarming motility, and pyocyanin overproduction.
200 luding biofilm formation, flagellum-mediated swarming motility, and type IV pilus-driven twitching.
201 hanced expression of the pilY1 gene inhibits swarming motility, and we identify residues in the putat
202 ulates the same phenotypes as BDSF including swarming motility, biofilm formation, and virulence.
203 riophage DMS3 inhibits biofilm formation and swarming motility, both important bacterial group behavi
204 e data, a DeltadipA mutant exhibited reduced swarming motility, increased initial attachment, and pol
205  production, high collagenase activity, high swarming motility, low resistance to chloramphenicol, an
206 dition, ppGpp deficiency resulted in loss of swarming motility, reduction of pyoverdine production, i
207 ve histidine kinase), and PAO1 DeltaalgR for swarming motility, rhamnolipid production, and rhlA tran
208 rsely controls biofilm formation and surface swarming motility, with high levels of this dinucleotide
209  regulate genes for biofilm formation or for swarming motility-the output phenotypes.
210 rexpression of MotA from a plasmid represses swarming motility.
211 PNPase mutant and also predicted a defect in swarming motility.
212 uired for the putrescine-dependent rescue of swarming motility.
213 ve over agar surfaces by flagellum-dependent swarming motility.
214 lus assembly machinery, in the repression of swarming motility.
215 lator activity have a commensurate effect on swarming motility.
216 uired for a form of surface migration called swarming motility.
217 oxidative stress, antibiotic resistance, and swarming motility.
218 -yet-unknown downstream mechanism to repress swarming motility.
219 rain had no effect on swimming but abolished swarming motility.
220 p behaviors, including biofilm formation and swarming motility.
221 system also contributes to the regulation of swarming motility.
222 hat YenS plays a direct, stimulatory role in swarming motility.
223  inverse regulation of biofilm formation and swarming motility.
224 ols mucoidy, while LadS negatively regulates swarming motility.
225 ss 3 promoters was found to be important for swarming motility.
226  aeruginosa, including biofilm formation and swarming motility.
227 ced biofilm formation but repressed swimming/swarming motility.
228 s complementary sense RNA levels and impacts swarming motility.
229 wosA allele resulted in a slight decrease in swarming motility.
230 de antibiotic sublancin and are defective in swarming motility.
231 e expression and FliZ's role in swimming and swarming motility.
232 32) of B. subtilis EF-P that is required for swarming motility.
233 olyl motifs present in proteins required for swarming motility.
234 ormation, quorum responses, development, and swarming motility.
235  as colony morphology, matrix formation, and swarming motility.
236 of 5-aminopentanonated EF-P is inhibitory to swarming motility.
237  has been shown to repress both swimming and swarming motility.
238 protein complexes necessary for swimming and swarming motility.
239 y of biological groups form a self-organized swarming motion at some point during their life spans, w
240  suppressors, inadvertent genetic linkage to swarming mutations, and auxotrophy.
241                                              Swarming not only allows bacteria to forage for food, bu
242 re systems in which both synchronization and swarming occur together.
243          We thus conclude that initiation of swarming occurs in response to specific cues and that ma
244 cterial flagellum, powering the swimming and swarming of many motile bacteria.
245                              Because initial swarming of neutrophils at the site of infection occurs
246 llular role of TFP during flagellar-mediated swarming of P. aeruginosa that does not require TFP exte
247                                              Swarming on agar to which chloramphenicol had been added
248 ine was found to be a strict requirement for swarming on normal swarm agar in addition to being a swa
249 histidine, malate, and DL-ornithine promoted swarming on several types of media without enhancing swi
250 cating that the surface translocation is not swarming or twitching motility.
251 ked for defects in swimming through liquids, swarming over surfaces, and transcriptional regulation.
252           Our simulations showed that TFP of swarming P. aeruginosa should be distributed all over th
253 mainly use computer simulations to study the swarming phenomena, this paper provides an analytical me
254 ed genes was altered consistent with the non-swarming phenotype observed for the mutant.
255 ng motility was established by analyzing the swarming phenotype of the PA3347 knock-out mutant and it
256 of the master regulators is important to the swarming phenotype perhaps at the level of controlling f
257 r exhibiting significant variation in social swarming phenotypes and five harboring large variation i
258                            Comparison of the swarming phenotypes of the single and double mutants and
259 rium with a complex life cycle that includes swarming, predation and fruiting body formation.
260 eactions exhibit collective behavior such as swarming, predator-prey interactions, and chemotaxis tha
261 es and nutrient requirements involved in the swarming process have been identified, few studies have
262                 It is puzzling however, that swarming-proficient and virulent strains of Vibrio parah
263               In these studies, we examine a swarming-proficient, virulent strain and identify an alt
264 es of nanomotors, which assemble in distinct swarming regions, is illustrated.
265 connected by a negative-feedback loop on the swarming regulon propagated by ExsA.
266                                              Swarming represents a special case of bacterial behavior
267                                 Furthermore, swarming repression by PilY1 specifically requires the d
268     We show that this HHQ- and PCA-dependent swarming repression is apparently independent of changes
269 ly but still functional in c-di-GMP-mediated swarming repression, indicating our ability to separate
270                                              Swarming requires that each cell periodically reverse it
271 ated during consolidation compared to during swarming, revealed that although these genes play a mino
272                    Furthermore, we show that swarming species do not form a monophyletic group and no
273 ies do not form a monophyletic group and non-swarming species that are closely related to locusts oft
274 their composition and remain in a collective swarming state or even differentiate among behavioral ph
275 croscopic colonial expansion, especially for swarming strains, but no measurements have been obtained
276                                              Swarming strategies are as diverse as the bacteria that
277 ageenan agar that may lay the foundation for swarming studies of snake-like, nonrod-shaped motile cel
278 ed (tan) colonies but still phase varied for swarming suggesting that pigmentation is not the cause o
279 mental conditions, selection for spontaneous swarming suppressors, inadvertent genetic linkage to swa
280 rve the swirling that is conspicuous in many swarming systems, probably due to increasingly long-live
281 ned syringafactin production exhibited broad swarming tendrils, while a syringafactin-producing strai
282 rain that overproduced HAA exhibited slender swarming tendrils.
283 vior-based computational model of M. xanthus swarming that allows the organization of cells to be com
284 ways, forms branched tendril patterns during swarming; this phenomena occurs only when bacteria produ
285                           Avaroferrin blocks swarming through its ability to bind iron in a form that
286 es have suggested that FliL is essential for swarming through its involvement in viscosity-dependent
287 n this region restored biofilm formation and swarming to DMS3 lysogenized strains.
288 duction of the capsule and the repression of swarming to elucidate the global scope of genes in the O
289 egulator or overexpression of flhDC restored swarming to the waaL mutant, despite the absence of O an
290 teria move in groups, in a mode described as swarming, to colonize surfaces and form biofilms to surv
291 wer cell density) between antagonist strains swarming toward each other.
292 pt, including swimming in aqueous media, and swarming, twitching and gliding on solid and semi-solid
293 ype IV pilus biogenesis, function to repress swarming via modulation of intracellular c-di-GMP levels
294 age series to capture the motion dynamics of swarming Vibrio alginolyticus at cellular resolution ove
295 in wosA-overexpressing strains, the onset of swarming was not altered.
296 understand the molecular signals controlling swarming, we isolated two bacterial strains from the sam
297 y to flagella-dependent migration modes like swarming, we show that this much faster "colony surfing"
298 ates, were tested for the ability to promote swarming when added to normally nonpermissive media.
299 al suspensions, granular materials and crowd swarming, where consequences may be dramatic.
300             Here, we focus on two aspects of swarming, which, if understood, hold the promise of reve

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